:root {
  /* Paradise palette (see /onyx/docker/*/theme/paradise.css); fonts stay Fraunces + Inter */
  color-scheme: dark;
  --ink: #e8e3e3;
  --ink-soft: #b8b0b0;
  --muted: #958d8d;
  --paper: #151515;
  --card: #1d1d1d;
  --raised: #242424;
  --line: #424242;
  --line-soft: #2e2e2e;
  --green: #8c977d;
  --link: #8da3b9;
  --link-hover: #aabdd0;
  --sun: #d9bc8c;
  --sun-hover: #e6cfa6;
  --red: #d18a8d;
  --night: #151515;
  --radius: 14px;
  --serif: "Fraunces", Georgia, serif;
  --sans: "Inter", system-ui, -apple-system, "Segoe UI", sans-serif;
}

* { box-sizing: border-box; }
html { scroll-behavior: smooth; scroll-padding-top: 72px; }
body { margin: 0; background: var(--paper); color: var(--ink); font: 17px/1.6 var(--sans); -webkit-font-smoothing: antialiased; }
img { display: block; max-width: 100%; }
a { color: var(--link); text-underline-offset: 3px; }
a:hover { color: var(--link-hover); }
h1, h2, h3 { font-family: var(--serif); line-height: 1.12; margin: 0; letter-spacing: -0.01em; }
h2 { font-size: clamp(2rem, 4vw, 2.9rem); margin-bottom: 2rem; }
h3 { font-size: 1.4rem; }
h4 { margin: 0; font-size: 1rem; }
.wrap { width: min(1160px, 100% - 40px); margin-inline: auto; }
.eyebrow { margin: 0 0 .6rem; font-size: .78rem; font-weight: 600; letter-spacing: .14em; text-transform: uppercase; color: var(--green); }
.muted { color: var(--muted); font-size: .95rem; }

/* Nav */
.nav { position: sticky; top: 0; z-index: 10; background: rgba(21, 21, 21, .9); backdrop-filter: blur(10px); }
.nav-row { display: flex; align-items: center; justify-content: space-between; gap: 1rem; min-height: 60px; }
.brand { color: var(--ink); text-decoration: none; font-family: var(--serif); font-weight: 700; font-size: 1.15rem; white-space: nowrap; }
.brand span { color: var(--sun); }
.nav nav { display: flex; gap: 1.4rem; }
.nav nav a { color: rgba(232,227,227,.82); text-decoration: none; font-size: .93rem; font-weight: 500; }
.nav nav a:hover { color: var(--ink); }

/* Hero */
.hero { position: relative; min-height: min(88vh, 820px); display: flex; align-items: flex-end; color: var(--ink); overflow: hidden; background: var(--night); }
.hero-bg { position: absolute; inset: 0; width: 100%; height: 100%; object-fit: cover; object-position: center 62%; }
.hero::after { content: ""; position: absolute; inset: 0; background: linear-gradient(180deg, rgba(21,21,21,.05) 0%, rgba(21,21,21,.4) 45%, rgba(21,21,21,.96) 100%); }
.hero-content { position: relative; z-index: 1; padding: 0 0 clamp(3rem, 8vw, 6rem); }
.hero .eyebrow { color: var(--sun); }
.hero h1 { font-size: clamp(2.4rem, 6vw, 4.6rem); line-height: 1.05; max-width: 17ch; text-wrap: balance; }
.lede { max-width: 40rem; font-size: clamp(1.05rem, 1.6vw, 1.25rem); color: rgba(232,227,227,.88); margin: 1.4rem 0 2rem; }
.actions { display: flex; flex-wrap: wrap; gap: .8rem; }
.btn { display: inline-block; padding: .85rem 1.5rem; border-radius: 999px; font-weight: 600; text-decoration: none; font-size: .98rem; }
.btn.primary { background: var(--sun); color: var(--night); }
.btn.primary:hover { background: var(--sun-hover); color: var(--night); }
.btn.ghost { border: 1.5px solid rgba(232,227,227,.6); color: var(--ink); }
.btn.ghost:hover { background: rgba(232,227,227,.12); color: var(--ink); }

/* Sections */
.section { padding: clamp(4rem, 9vw, 7rem) 0; }

/* Research cards */
.cards { display: grid; grid-template-columns: repeat(3, 1fr); gap: 1.4rem; }
.card { background: var(--card); border: 1px solid var(--line-soft); border-radius: var(--radius); overflow: hidden; display: flex; flex-direction: column; }
.card img { width: 100%; aspect-ratio: 16 / 10; object-fit: cover; }
.card-body { padding: 1.4rem 1.5rem 1.6rem; display: flex; flex-direction: column; gap: .7rem; flex: 1; }
.card-body p { margin: 0; color: var(--ink-soft); font-size: .98rem; }
.card .cite { margin-top: auto; padding-top: .4rem; font-size: .9rem; font-weight: 500; }
.credit { font-size: .75rem !important; color: var(--muted) !important; }
.credit a { color: inherit; }
.card.feature { grid-column: 1 / -1; display: grid; grid-template-columns: 1.25fr 1fr; }
.card.feature img { height: 100%; aspect-ratio: auto; min-height: 320px; }
.card.feature .card-body { padding: 2.2rem; justify-content: center; }
.card.feature h3 { font-size: 2rem; }
.card.feature p { font-size: 1.05rem; }
.card.feature .cite { margin-top: .4rem; }

.underway { margin-top: 3rem; background: var(--card); border: 1px solid var(--line-soft); border-radius: var(--radius); padding: 2rem 2.2rem; }
.underway h3 { margin-bottom: 1rem; }
.underway ul { margin: 0; padding: 0; list-style: none; display: grid; grid-template-columns: repeat(2, 1fr); gap: .9rem 2.4rem; }
.underway li { color: var(--ink-soft); font-size: .96rem; padding-left: 1.1rem; position: relative; }
.underway li::before { content: ""; position: absolute; left: 0; top: .62em; width: 7px; height: 7px; border-radius: 50%; background: var(--green); }
.underway strong { color: var(--ink); }
.tag { display: inline-block; font-size: .7rem; font-weight: 600; text-transform: uppercase; letter-spacing: .08em; background: rgba(209,138,141,.15); color: var(--red); border-radius: 4px; padding: .05rem .4rem; margin-right: .2rem; vertical-align: 1px; }

/* Tools */
.tools { background: var(--card); color: var(--ink); border-block: 1px solid var(--line-soft); }
.tools .eyebrow { color: var(--sun); }
.tool-grid { display: grid; grid-template-columns: repeat(3, 1fr); gap: 1rem; }
.tool { display: block; padding: 1.4rem 1.5rem; border-radius: var(--radius); background: var(--raised); border: 1px solid var(--line-soft); color: var(--ink); text-decoration: none; transition: background .15s, transform .15s; }
.tool:hover { background: #2a2a2a; border-color: var(--line); color: var(--ink); transform: translateY(-2px); }
.tool h3 { font-size: 1.2rem; margin-bottom: .4rem; }
.tool h3::after { content: " →"; color: var(--sun); }
.tool p { margin: 0; color: rgba(232,227,227,.75); font-size: .93rem; }

/* Publications */
.pubs { display: grid; grid-template-columns: 1fr 2.6fr; gap: 3rem; }
.pub-list { list-style: none; margin: 0; padding: 0; }
.pub-list li { display: grid; grid-template-columns: 3.5rem 1fr; gap: 1rem; padding: 1rem 0; border-top: 1px solid var(--line-soft); }
.pub-list span { font-weight: 600; color: var(--sun); }
.pub-list p { margin: 0; font-size: .95rem; color: var(--ink-soft); }
.pub-list a { color: var(--ink); text-decoration: none; font-weight: 500; }
.pub-list a:hover { color: var(--link); text-decoration: underline; }

/* People */
.people { background: var(--card); border-top: 1px solid var(--line-soft); }
.pi { display: grid; grid-template-columns: 260px 1fr; gap: 2.4rem; align-items: start; background: var(--raised); border-radius: var(--radius); border: 1px solid var(--line-soft); padding: 2rem; }
.pi img { width: 100%; aspect-ratio: 1; object-fit: cover; border-radius: 12px; }
.pi h3 { font-size: 1.8rem; }
.pi .role { font-weight: 600; color: var(--sun); margin: .5rem 0 .8rem; }
.roles { margin: 0 0 1rem; padding-left: 1.1rem; color: var(--ink-soft); font-size: .96rem; }
.pi p { color: var(--ink-soft); }
.links { display: flex; flex-wrap: wrap; gap: 1.2rem; font-weight: 600; font-size: .95rem; }
.students-head { margin-top: 3.5rem; }
.students { display: grid; grid-template-columns: repeat(3, 1fr); gap: 1rem; margin-top: 1.4rem; }
.students article { display: flex; gap: .9rem; align-items: center; background: var(--raised); border: 1px solid var(--line-soft); border-radius: 12px; padding: .8rem; }
.students img, .initials { width: 64px; height: 64px; flex: none; border-radius: 50%; object-fit: cover; }
.initials { display: grid; place-items: center; background: var(--green); color: var(--night); font-weight: 600; letter-spacing: .04em; }
.students p { margin: .15rem 0 0; font-size: .86rem; color: var(--muted); line-height: 1.4; }

/* Contact + footer */
.contact { background: var(--night); color: var(--ink); padding: clamp(3.5rem, 7vw, 5rem) 0; }
.contact .eyebrow { color: var(--sun); }
.contact h2 { margin-bottom: .6rem; }
.contact p { color: rgba(232,227,227,.75); margin: 0; }
.contact-card { display: flex; flex-wrap: wrap; justify-content: space-between; align-items: center; gap: 2rem; }
.contact-links { display: flex; flex-direction: column; gap: .6rem; }
.contact-links a { color: var(--ink); font-size: 1.1rem; font-weight: 500; }
.contact-links a:hover { color: var(--sun); }
.footer { background: var(--night); border-top: 1px solid rgba(232,227,227,.1); color: rgba(232,227,227,.6); font-size: .88rem; }
.footer-row { display: flex; flex-wrap: wrap; justify-content: space-between; gap: 1rem; padding: 1.4rem 0; }
.footer p { margin: 0; }
.social { display: flex; gap: 1.2rem; }
.social a { color: rgba(232,227,227,.8); text-decoration: none; }
.social a:hover { color: var(--sun); }

/* Responsive */
@media (max-width: 960px) {
  .cards, .tool-grid, .students { grid-template-columns: repeat(2, 1fr); }
  .card.feature { grid-template-columns: 1fr; }
  .card.feature img { min-height: 0; aspect-ratio: 16 / 9; }
  .pubs { grid-template-columns: 1fr; gap: 0; }
}
@media (max-width: 680px) {
  body { font-size: 16px; }
  .wrap { width: calc(100% - 32px); }
  .nav-row { flex-direction: column; align-items: flex-start; gap: .2rem; padding: .7rem 0 .5rem; }
  .nav nav { gap: 1.1rem; overflow-x: auto; max-width: 100%; }
  .nav nav a { font-size: .85rem; white-space: nowrap; }
  html { scroll-padding-top: 96px; }
  .cards, .tool-grid, .students, .underway ul { grid-template-columns: 1fr; }
  .card.feature .card-body { padding: 1.5rem; }
  .pi { grid-template-columns: 1fr; padding: 1.4rem; }
  .pi img { max-width: 220px; }
  .underway { padding: 1.5rem; }
}

.card img.top { object-position: center 12%; }

/* Applied ecology */
.section-lede { max-width: 44rem; color: var(--ink-soft); margin: -1.2rem 0 2rem; font-size: 1.05rem; }
.applied-grid { display: grid; grid-template-columns: repeat(4, 1fr); gap: 1.2rem; }
@media (max-width: 960px) { .applied-grid { grid-template-columns: repeat(2, 1fr); } }
.applied-grid article { background: var(--card); border: 1px solid var(--line-soft); border-left: 3px solid var(--sun); border-radius: var(--radius); padding: 1.5rem 1.6rem; display: flex; flex-direction: column; gap: .6rem; }
.applied-grid h3 { font-size: 1.3rem; }
.applied-grid p { margin: 0; color: var(--ink-soft); font-size: .98rem; }
.applied-grid .cite { margin-top: auto; padding-top: .3rem; font-size: .88rem; font-weight: 500; color: var(--muted); }
@media (max-width: 680px) { .applied-grid { grid-template-columns: 1fr; } }

.press { margin: 0; font-size: .9rem !important; color: var(--muted) !important; }
.press a { color: var(--sun); }
.press a:hover { color: var(--sun-hover); }

/* About / expertise */
.about { background: var(--card); border-bottom: 1px solid var(--line-soft); }
.about-grid { display: grid; grid-template-columns: 1.4fr 1fr; gap: 3rem; align-items: start; }
.about h2 { margin-bottom: 1rem; }
.about-lede { color: var(--ink-soft); font-size: 1.05rem; margin: 0 0 1.8rem; }
.about .btn.ghost { border-color: var(--line); color: var(--ink); }
.expertise { background: var(--raised); border: 1px solid var(--line-soft); border-radius: var(--radius); padding: 1.6rem; }
.expertise h3 { margin-bottom: 1rem; font-size: 1.2rem; }
.chips { list-style: none; margin: 0; padding: 0; display: flex; flex-wrap: wrap; gap: .5rem; }
.chips li { font-size: .88rem; padding: .35rem .75rem; border-radius: 999px; background: rgba(140,151,125,.14); border: 1px solid rgba(140,151,125,.35); color: var(--ink); }
.pub-list a.data { font-size: .82rem; font-weight: 600; color: var(--green); margin-left: .3rem; white-space: nowrap; }
.pub-list a.data:hover { color: var(--sun); }
@media (max-width: 960px) { .about-grid { grid-template-columns: 1fr; gap: 2rem; } }